The Penguin update doesn't seem to be related at all to on-site factors according to experts, so you wouldn't need to change much there. It seems to be penalizing over-optimization of off-page factoes, especially anchor text "spamming". That is, is your site is "buybluewidgets.com" and you have built 1000 backlinks that all link to your site using "buy blue widgets", that seems to trigger a penguin penalty. I'd diversify the backlink profile. Create lots of backlinks using naked links, that is using buybluewidgets.com or the www. version, or the http:// version as the anchor text to your site. Also add some variation, using your name or pen-name as the anchor text, using LSI keywords, using general keywords like "click here", etc. As I said, I'm not an expert, but that's what most experts are saying, use your target keyword as the anchor text to your site only in 30 to 50% of the cases, and vary the rest of your backlink profile.
Subscribed to said above + unique content. I can see that they are focusing on unique content, no matter what quality.
After Penguin update, You should take care of your Social Media Presence and also the link neighbors.. i.e dont build links from Spam pages or pages which have lot of Out Bound Links. Contextual Links are working great (as always). Happy to answer any more questions
You can move on like how you did before. All you have to know is "Never Spam" and "not to over optimize" by building too many backlinks via automated tools, link building services etc. Do it manually and naturally, produce quality, relevant and original contents and use the off-page techniques to distribute your content and then you can get some valuable backlinks which are really safe and keep you away from the Google's punishment.
as always said by google, quality matters but not quantity.., building links without quality and in a bulk with single anchor text were hit.., rotate ur anchor text frequently, do quality submissions, then ur doesn't effects even there are huge updates..,
Check out whether your anchor text is not of a type that is penalized by Google. Use "click here", or "read more" kind of anchor text to avoid Google penalty.
Agree with Coro. There's a good article on Webpronews (don't know the url) where they talk about the impact on making your site being shared a lot on social networks and its correlation with Google ranks.
Exact match anchor text is now bad news, if you have a brand, use that more in your anchor text, much safer, keeps you on the right side of google.
Content is the king for good visitors , so enrich your site with quality content and build link with social media .